Barkworth

Mary E. Barkworth

Position

Director, Intermountain Herbarium, Utah State University

Associate Professor (retired), Utah State University

Contact information

Mail:     Department of
Biology, Utah State
University, Logan, Utah 84322-5305

Email:   mary@biology.usu.edu

Voice:  (435)-797-1584           FAX:    (435)-797-1575

History

Education

B. Sc.               1961    University of British
Columbia (Canada);
Majors: Mathematics & Physics

PBTC              1965    [=Professional Basic Teaching Certificate]

                                    Secondary
School Science, Province
of British Columbia

M. Ed.                         1970    Western Washington
State College; General
Science Education

Ph.D.               1975    Washington
State University;
Major: Botany, Minor: Biochemistry

Employment

Teacher                        9/62-6/68         Teacher, Junior high school in British Columbia

Grad. Asst.                   9/68-8/69         Department of Physics, Western Washington
State College

Teach. Asst.                 9/69-6/72         Department of Botany, Washington State
University

Curatl. Ast.                   7/72-8/75         Marion Ownbey Herbarium, Washington State
University

Research Sci.               7/75-3/79         Biosystematics Research Inst., Agriculture Canada

Asst. Prof.                    3/79-6/85         Biology
Department, Utah
State University

Director                        7/80-6/82         Intermountain Herbarium, Utah State
University

Assoc. Prof.                 7/85-6/12         Biology
Department, Utah
State University       

Herbarium Director       1/88-Now        Intermountain
Herbarium, Utah State University

Publications

Student names are underlined.

Refereed publications

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  1. Barkworth, M.E.
    and Z. Murrell. (2012 - accepted). The US Virtual Herbarium Initiative:
    Building a national resource from the ground up. Zookeys.
  2. Olonova,
    M.V, M.E. Barkworth, S.V. Pulkina
    and W-L Chen. 2012. Morphological variation in populations of the Poa orinosa complex (Poaceae: Poeae) in northeastern China. Nordic Journal of Botany
    29:762-772. DOI: 10.1111/j.1756-1051.2011.01277.x
  3. Barkworth, M.E.
    and S.W.L. Jacobs. 2011. The Triticeae
    in Australasia. Telopea 13(1-2): 37-56.
  4. Mercader,
    J., F. Studillo, M.E. Barkworth, T.
    Bennett, C. Esselmont, R. Kinyanjui, D. L. Grossman, S. Simpson, and D. Walde.
    2010. Poaceae phytoliths from the
    Niassa Rift, Mozambique. Journal of Archeological
    Science 37: 1953-1967.
  5. Barkworth, M.E.,
    R. Cutler, J. Rollo, S.W.L. Jacobs, and A. Rashid. 2009. Morphological
    identification of genomic genera in the perennial Triticeae (Poaceae).
    Breeding Science (Special Edition) 59: 561-570.
  6. Barkworth, M.E.,
    S.W.L. Jacobs, and H.Q. Zhang. 2009. Connorochloa,
    a new genus in the Triticeae. Breeding Science (Special Edition) 59: 685-686.
  7. Jacobs,
    S.W.L. and M.E. Barkworth. 2009. A
    new species of Elymus (Gramineae, Triticeae) from eastern Australia. Novon 19: 168–171.
  8. Barkworth, M.E.,
    M. O. Arriaga, J.F. Smith, S.W.L. Jacobs, J.Valdes-Reyna, and B. S.Bushman.
    2008. Molecules and morphology in South American Stipeae (Poaceae).
    Systematic Botany 33: 719–731.
  9. Whipple,
    I., M.E. Barkworth, and B.S. Bushman. 2007.
    Molecular insights into the taxonomy of Glyceria
    (Poaceae: Meliceae) in North America.
    American Journal of Botany 94(4):551–557.

10.  Barkworth, M.E. 2006. A new hybrid genus and 12 new
combinations in North American grasses. Sida 22:495-501.

11.  Barkworth, M.E.,
L.K. Anderton, J. McGrew, and D.E. Giblin. 2006. Geography and morphology of
the Bromus carinatus (Poaceae: Bromeae) complex. Madroño 53:235–245.

12.  Arriaga,
M.O. and M.E. Barkworth. 2006. Amelichloa: a new genus in the Stipeae
(Poaceae). Sida 22:145–149.

13.  F.M.
Vásquez and M.E. Barkworth. 2004.
Resurrection and Emendation of Macrochloa
(Gramineae: Stipeae). Botanical Journal of the Linnaean Society 144:483–495.

14.  Gandhi, K.
and M.E. Barkworth. 2003. Nomenclatural
and Taxonomic Review of Knotroot Bristle Grass (Setaria parviflora, Gramineae).
Rhodora 197–204.

15.  Gandhi, K.
and M.E. Barkworth. 2003. Proposal
to conserve the name Munroa (Gramineae) with that spelling. Taxon
52:137-138.

16. 
Valdés-Reyna, J. and M.E. Barkworth. 2002. Stipeae. Flora de Veracruz, Fasc. 127. Poaceae II, Pooideae: Tribu Stipeae.
Instituto de Ecología, A.C., Xalapa, Veracruz, México.

17.  Barkworth, M.E.
and S.W.L. Jacobs. 2001. Valuable research or short stories: what makes the difference?
Hereditas 135:263–270.

18.  Barkworth, M.E.
and M.A. Torres. 2001. Distribution and diagnostic characters of Nassella (Poaceae: Stipeae). Taxon
50:439-468.

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19.  Barkworth, M.E.,
R.L. Burnham, and L.E. Talbert. 
1996.  Elymus calderi: a new species in the Triticeae (Poaceae).  Systematic Botany 21:349-354.

20.  Jacobs,
S.W.L., J. Everett, and M.E. Barkworth.  1995. 
Clarification of morphological terms used in the Stipeae (Gramineae) and a
reassessment of Stipa in Australia.
Taxon 43: 33‑42.

21.  Valdes
Reyna, J. and M.E. Barkworth.  1994.  El genero Nassella
(Poaceae: Stipeae) en Mexico. Acta Botanica Mexicana, 26: 63‑75.

22.  Barkworth, M.E.  1993. Proposal to conserve the type of Stipa robusta (Poaceae) under Article
69.3. Taxon 42: 711‑713.

23.  Jarvie,
J.K. and Barkworth, M.E. 
1992.  Morphological variation and
genome constitution in some perennial Triticeae.
Botanical Journal of Linnean Society, 108: 167‑180.

24.  Jarvie,
J.K. and M.E. Barkworth.  1992.  Anatomical variation in some perennial Triticeae.  Botanical Journal of Linnaean Society, 108:
287‑301.

25.  Barkworth, M.E.  1990. Nassella
(Gramineae: Stipeae): Revised interpretation and nomenclatural changes. Taxon,
39: 597‑614.

26.  Jarvie,
J.K. and M.E. Barkworth.  1990. 
Isozyme similarity in Thinopyrum
and its relatives (Triticeae: Gramineae)n. Genome, 31: 885‑891.

27.  Barkworth, M.E.,
J. Valdes‑Reyna and R.Q. Landers, Jr. 
1989.  Stipa clandestina: New weed threat on southwestern rangelands.  Weed Techn., 3: 699‑702.

28.  Fermanian,
T., M.E. Barkworth, and H. Liu.  1989. 
Trained and untrained Individuals' ability to identify morphological
characters of immature grasses. Agronomy Journal, 81: 918‑922.

29.  Barkworth, M.E.  1988. 
New taxa in Piptochaetium (Stipeae, Gramineae) from Mexico.  Syst. Bot., 13: 196‑201.

30.  Barkworth, M.E.  1986. Piptochaetium
(Gramineae, Stipeae) in North and Mesoamerica:
taxonomic and distributional observations. 
Brenesia, 25‑26: 169‑178.

31.  Barkworth, M.E.
and D.R. Dewey.  1985.  Genomically based genera in the perennial Triticeae of North
America: identification and limits.  American Journal of Botany, 72: 767‑776.

32.  Barkworth, M.E.
and R.J . Atkins.  1984.  Leymus
Hochst. (Gramineae: Triticeae) in North
America: taxonomy and distribution.  American Journal of Botany, 71: 609‑625.

33.  Barkworth, M.E.
and J. Linman.  1984. Stipa lemmonii (Gramineae: Stipeae): A
taxonomic and distributional study. Madroño, 31: 48‑56.

34.  Neely, E.E.
and M.E. Barkworth.  1984. 
Vegetation on soils derived from dolomite and quartzite in the Bear
River Range, Utah: a comparative study. 
Bull. Torrey Bot. Club., 111: 179‑192.

35.  Atkins,
R.J., M.E. Barkworth and D.R.
Dewey.  1984.  A taxonomic study of Leymus ambiguus and L.
salinus
(Poaceae: Triticeae). Syst. Bot., 9: 279‑294.

36.  Barkworth, M.E.  1983.  
Ptilagrostis Griseb. in North America and its relationship to other taxa in the Stipeae (Gramineae). Syst. Bot., 8: 395‑415.

37.  Barkworth, M.E.,
D.R. Dewey and R.J. Atkins.  1983.  New generic concepts in the Triticeae of the Intermountain Region:
keys and comments. Great Basin Naturalist, 43:
561‑572.

38.  Barkworth, M.E.  1982. 
The identity of Stipa columbiana
Macoun and Stipa viridula var. minor Vasey: a  rebuttal. Taxon, 31: 290‑299.

39.  Barkworth, M.E.  1982. 
Embryological characters and the taxonomy of the Stipeae (Gramineae). Taxon, 31: 233‑243.

40.  Barkworth, M.E.  1981. 
Foliar epidermes and the taxonomy of North American Stipeae. Syst. Bot., 6: 136‑152.

41.  Barkworth, M.E.  1979.  Proposal to Reject Stipa columbiana Macoun and nomenclatural changes affecting three
western North American species of Stipa.
Taxon, 28: 621‑625.

42.  Barkworth, M.E.,
J. McNeill, and J. Maze.  1979.   A taxonomic study of Stipa nelsonii (Poaceae)
with a key distinguishing it from related taxa in western North
America. Canadian Journal of Botany, 57: 2539‑2553.

43.  Barkworth, M.E.  1978. 
A taxonomic study of the large-glumed species of Stipa (Gramineae) Occurring in Canada. Canadian Journal of Botany,
56: 606‑625.

44.  Barkworth, M.E.  1977. 
Intraspecific variation in Brodiaea
douglasii
Watson (Liliaceae). Northwest Science, 51: 79‑90.

45.  Barkworth, M.E.  1973.  Impatiens parviflora in British Columbia.
Madrono, 2: 24.

Books and proceedings edited

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  1. Anderton, L.K. and M.E.
    Barkworth
    (Eds.). 2009. Grasses of the Intermountain Region. Intermountain
    Herbarium, Utah State University, Logan, Utah. 559 pp. [Derived from 3 and 5]
  2. Barkworth, M.E.,
    L.K. Anderton, K.M. Capels, S. Long, and M.B. Piep.  (Editors). 2007. Manual of grasses for North
    America. Utah State University Press. 636 pp. [Derived from 3 and 5]
  3. Barkworth, M.E.,
    K.M. Capels, S. Long, L.K. Anderton, and M.B. Piep, editors. 2007. Flora of
    North America, volume 24. Poaceae,
    part 1. Oxford University Press, New York. 911 pp.
  4. Holubec, V., M.E.
    Barkworth
    , and R. von Bothmer (Eds). 2005. Proceedings of the Fifth
    International Triticeae Symposium, Prague, June 6-10, 2005. Czech J. Genet.
    Plant. Breed. 41 (Special Issue). 344 pp.
  5. Barkworth, M.E.,
    K.M. Capels, S. Long, M.B. Piep (editors). 2003. Flora of North America, volume
    25. Poaceae, part 2. Oxford
    University Press. 793 pp.

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  1. Soderstrom, T.R., K. Hilu, C.S. Campbell, and M.E. Barkworth, editors. 1987. Grass
    systematics and evolution. Smithsonian Institution Press, Washington, D.C. 474
    pp.

Floristic treatments.

These were all reviewed by other taxonomists before
acceptance, including those in volumes for which I was an editor. They are
grouped by their publication and by authorship.

  1. I.       Treatments
    significantly modified for Anderton, L.K. and M.E. Barkworth (Eds.). 2009. Grasses of the Intermountain Region.
    Intermountain Herbarium, Utah State University, Logan, Utah. 559 pp.

Trisetum (p.); Panicum (p. ).

II. Treatments in Barkworth,
M.E.
, K.M. Capels, S. Long, L.K. Anderton, and M.B. Piep, editors. 2007.
Flora of North America, volume 24. Poaceae, part 1. Oxford University Press,
New York.

Barkworth, M.E. 2007. Key to tribes (p. 7–10), Olyreae (p. 29), Lithachne
(p. 30), Ehrharteae, Ehrharta (p. 33–36), Meliceae (p. 67–68), Glyceria (p. 68–88), Melica (p. 88–102), Stipeae (p. 109–111), Achnatherum
(p. 114–142), Ptilagrostis (p.
143–144), Piptatherum (p. 144–151), Stipa (p. 154–156), Hesperostipa (p. 157–161), Piptochaetium
(p. 161–167), Oryzopsis (p. 167–168),
×Achnella  (p. 169–170), Nassella (p. 170–177), Triticeae
(p. 238­–240), Agropyron (p.
277–279), ×Elyhordeum (p.
283–287),  Pascopyrum (p. 350–351), Leymus
(p. 353–369), Thinopyrum (p.
373–378), Poeae (p. 378–388), Deschampsia (p. 624–633), Polypogon (p. 662–668), ×Agropogon (P. 608), Phleum (p. 670–675), Sesleria
(p. 678­–681), Scolochloa (p.
732–734),  Phalaris (p. 764–773), Ammophila
(p. 776–778)

Barkworth, M.E., J.J.N. Campbell, and B. Salomon. 2007. Elymus. Pp. 288–348.

Barkworth, M.E. and E.E. Terrell. 2007. Oryza. Pp. 37–41.

Allred, K.W. and M.E. Barkworth. 2007. Anthoxanthum, Pp. 758–764.

Stapleton, C.M.A. and M.E. Barkworth. 2007. Phyllostachys. Pp. 25–27.

II. Treatments in Barkworth, M.E.,
K.M. Capels, S. Long, M.B. Piep (editors). 2003. Flora of North America, volume
25. Poaceae, part 2. Oxford University Press.

Barkworth, M.E. 2003.  Hakonechloa (p. 8), Cynodonteae (p. 14–21), Distichlis (p. 25), Aeluropus (p. 25–27), Dinebra (p. 63–64), Cladoraphis (p. 105), Chloris (p. 204–218), Enteropogon (p. 224), Trichoris (p. 225–227), Ctenium (p. 232–234), Cynodon (p. 235–240), Spartina (p. 240–250), Opizia (p. 269), Cathestecum (p. 272–273), Aegopogon (p. 273–274), Hilaria (p. 274–278), Paniceae (p. 353–358), Reimarochloa (p. 599–601), Andropogoneae (p. 602–608), Miscanthus (p. 616–618), Polytrias (p. 623), Sorghum (p. 626–630), Dichanthium (p. 637–638), Cymbopogon (p. 664–666), Hyparrhenia (p. 678–679), Heteropogon (p. 680–681), Themeda (p. 682–684, Elionurus (p. 684–685). Tripsacum (p. 693–696).

III. Treatment in COTECA, Las Gramíneas de México, Mexico City, México.

Valdés-Reyna, J. and M.E. Barkworth. 1999. Piptochaetium, pp. 339–352.

IV.       Treatments in J.C. Hickman [Ed.], The Jepson Manual of the Higher Plants of
California. University of California Press, Berkeley.  Pp. 1294.

Barkworth, M.E.  1993.  Agropyron
(p. 1252–1256), Elymus (p. 1253‑1256),
Glyceria (p. 1262‑1263), Hesperostipa (p. 1263), Hordeum  (p. 1264‑1266), Leymus ((p. 1268‑1270), Melica
((p. 1271‑1272), Nassella (p. 1274‑1276),
Pascopyrum ((p. 1280), Piptatherum (p. 1282‑1283), Piptochaetium((p. 1283), Pleuropogon (p. 1283‑1284), Pseudoroegneria (p. 1290), Ptilagrostis
(p. 1292), Secale(p. 1294) in

Jarvie, J.K. and M.E. Barkworth.  1993.  Elytrigia. In J.C. Hickman [Ed.], The
Jepson Manual of the Higher Plants of California. University of California
Press, Berkeley.  Pp. 1256‑1257.

V.
Barkworth, M.E.
  1980. Poaceae
pp. 310-376 in R.J. Shaw. 1980. Vascular plants of Northern Utah. Utah
University Press, Logan, Utah.

Chapters in Books

These were all reviewed before acceptance, including those
in volumes for which I was an editor.

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46.  Barkworth, M.E. and R. von Bothmer.
2009. Scientific names in the Triticeae.
Pp. 3–30 in C. Feuillet and G. J. Muehlbauer (Eds.), Genetics and Genomics of
the Triticeae, volume 7 in “Plant
Genetics and Genomics” Series Editor R. Jorgensen Springer Verlag.

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47.  Barkworth, M.E.  2000. 
Changing perceptions of the Triticeae.  Pp. 110-120 in S.W.L. Jacobs and J. Everett
(Editors), Proceedings of the Third International Symposium on Grass
Systematics and Evolution. CSIRO, Canberra, Australia.

48.  2.   Barkworth,
M.E.
and K.M. Capels.  2000.  The Poaceae
in North America: A Geographic Perspective. Pp. 327-346 in S.W.L. Jacobs and J.
Everett (Editors), Proceedings of the Third International Symposium on Grass
Systematics and Evolution. CSIRO, Canberra, Australia.

49.  3.   Jacobs, S.W.L., J. Everett, M.E. Barkworth, and C. Hsiao. 2000.
Relationships within the Stipoid grasses (Gramineae). Pp. 75-82 in S.W.L.
Jacobs and J. Everett (Editors), Proceedings of the Third International
Symposium on Grass Systematics and Evolution. CSIRO, Canberra, Australia.

50.  4.   Barkworth,
M.E.
and J. Everett. 1987. Evolution in the Stipeae: Identification and relationships of its monophyletic taxa.
Pp. 231-264 in Soderstrom, Hilu, Campbell and Barkworth (eds)., Grass systematics and evolution. Smithsonian
Institution Press, Washington, D.C.

Non-refereed publications – Web-based

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  1. Barkworth, M.E 2012. SYMBIOTA: Open
    Source Software for Developing Specimen-based Floristic Resources for Research
    and Education. (Presentation recorded and available at http://utahstatelive.wimba.com/launcher.cgi?room=_user_A00015751_class__2012_0202_0913_07

52.  Barkworth, M.E. 2012. Interactive
multi-access key to the spore-bearing vascular plant genera in the
Intermountain Region. See http://herbarium.usu.edy/keys

53.  Barkworth, M.E. 2011. Illustrated key
to the genera of Utah’s mosses based on the treatment by Seville Flowers.
Downloadable pdf. See http://herbarium.usu.edy/keys

54.  Barkworth, M.E. 2011. Interactive
dichotomous keys to the families of lycopods, monilophytes, and monocots that
reflect current taxonomic treatments. Also some keys for the genera in these
groups. http://utc.usu.edu/keys Created
using Lucid Phoenix®. (Centre for Biological Information Technology).

55.  Barkworth, M.E. 2009. Multi-access key
to Poa in the Intermountain Region,
based on the treatment by R.J. Soreng in Barkworth,
M.E.,
K.M. Capels, S. Long, L.K. Anderton, and M.B. Piep (editors). 2007. Flora of North
America
, volume 24. Poaceae, part 1. Oxford University Press, New
York. http://utc.usu.edu/keys/IMRPoa/.
Created using Lucid 3 (Centre for Biological Information Technology).

56.  Anderton,
L.K. and M.E. Barkworth. 2009.
Multi-access key to Festuca in the
Intermountain Region, based on the treatment by S.J. Darbyshire and L.
Pavlick  in Barkworth, M.E., K.M. Capels, S. Long, L.K. Anderton, and
M.B. Piep (editors). 2007. Flora
of North America
, volume 24. Poaceae, part 1. Oxford University
Press, New York. http://utc.usu.edu/keys/IMRFestuca/.
Created using Lucid 3 (Centre for Biological Information Technology).

57.  Barkworth, M.E. 2009. Dichotomous key
to the genera of the Triticeae.
Created using Phoenix (Centre for Biological Information Technology).
http://herbarium.usu.edu/Triticeae/

DichotKeys/Genera%20of%20the%20Triticeae.html

58.  Barkworth, M.E. 2009. Descriptions of
the genera of the Triticeae. Created
using Fact Sheet Fusion (Centre for Biological Information Technology).
http://herbarium.usu.edu/Triticeae/

Genera/index.htm

59.  Barkworth, M.E. 2009. Multi-access key
to the genera of the Triticeae. Created
using Lucid 3.5 (Centre for Biological Information Technology).
http://herbarium.usu.edu/Triticeae/

Keys/Genera/Genera%20of%20the%20Triticeae.html

60.  Barkworth, M.E. 2009. Dichotomous key
to the Australasian genera of the Triticeae.
Created using Phoenix (Centre for Biological Information Technology). http://herbarium.usu.edu/Triticeae/Keys/Australasian%20Triticeae.html

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61.  Barkworth, M.E. 2009. Descriptions of
the species of Anthosachne (Poaceae: Triticeae). Created using
Fact Sheet Fusion (Centre for Biological Information Technology). http://herbarium.usu.edu/Triticeae/Anthosachne/index.htm

In addition, I have placed numerous
notes relating to my teaching on the web. These are updated each year I teach a
class. There are, however, many pages that I have not revised.

Non-refereed publications –Print

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62.  Barkworth, M.E. 2012. Opening the region’s
herbaria. Sego Lily.

63.  Barkworth, M.E. and R. von Bothmer.
2005. Twenty-one years later: the impact of Löve and Dewey’s genomic
classification proposal. In Proc. Fifth International Triticeae Symposium, Prague, June 6–10,
2005. Czech J. Genet. Plant. Breed. 41 (Special Issue): 3–9.

64.  Barkworth, M.E. and J. McGrew.
2005. Combining herbarium data with spatial data: potential benefits, new
needs. In Proc. Fifth International Triticeae
Symposium, Prague, June 6–10, 2005. Czech J. Genet. Plant. Breed. 41 (Special
Issue): 59–64.

65.  Barkworth, M.E., J. McGrew, and
G. Zhu. 2005. The Triticeae in China. In Proc. Fifth
International Triticeae Symposium,
Prague, June 6–10, 2005. Czech J. Genet. Plant. Breed. 41 (Special Issue):
65–69.

66.  Barkworth, M.E. and S.-J. Lim. 2005. A Triticeae Website: report of discussion.
In Proc. Fifth International Triticeae
Symposium, Prague, June 6–10, 2005. Czech J. Genet. Plant. Breed. 41 (Special
Issue): 220–223.

67.  Bothmer, R.
von, M.E. Barkworth, and V. Holubec.
2005. What are the Triticeae and why a Triticeae Symposium? Proceedings of the Fifth International Triticeae Symposium, Prague, June 6-10,
2005. Czech J. Genet. Plant. Breed. 41 (Special Issue). 41 (Special Issue):
1–2.

68. 
Barkworth,
M.E.
and S.W.L. Jacobs. 2002. Voucher specimens: a critical element in a
scientific study.  Pp. 112-118 in P.
Hernández, M.T. Moreno, J.I. Cubero, and A. Martin (Editors),  Proceedings of the Fourth International Triticeae Symposium. Junta de Andalucia, Consejería de Agricultura y
Pesca, Córdoba, Spain.

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69.  Barkworth, M.E.  1997.  Elymus
stebbinsii
: Taxonomy, nomenclature, and distribution.  Phytologia 83:345-365.

70.  Carlson,
J.R. and M.E. Barkworth.  1997.  Elymus
wawawaiensis
: a species hitherto confused with Pseudoroegneria spicata (Triticeae, Poaceae).  Phytologia 83:312-330.

71.  Barkworth, M.E.  1997. 
Taxonomic and nomenclatural comments on the Triticeae in North America. 
Phytologia 83:302-311.

72.  McNeill, J.
and M.E. Barkworth.  1996. 
Rules for Botanical Nomenclature.  
Pp.21–26 in F. Salisbury,
(ed.) Units, symbols, and terminology for plant physiology : a reference for
presentation of research results in the plant sciences. Oxford University
Press, New York.

73.  Barkworth, M.E.  1993. North American Stipeae (Gramineae): Taxonomic changes and other comments.  Phytologia, 74: 1‑25.

74.  Barkworth, M.E., M.A. Kobler, T.S.
Stockton, E.E. Neeley and D. Henderson. 
1982. Woody plants of campus. Logan: Utah State University.  Extension Service.

75.  Barkworth, M.E.  1982. 
Bur Buttercup: A Weedy Immigrant. Utah Science, Spring: 3‑4.

76.  R.J. Shaw
and M.E. Barkworth.  1982. 
USU's Plant Photographic Library. Utah Science, 43: 26‑30.

Reports and newsletters

  1. Ayed, G. and M.E.
    Barkworth
    .  1988.  Report of the Review Team on Training
    Programs, International Board of Plant Genetic Resources. Submitted to the
    International Board of Plant Genetic Resources.

77.  Barkworth, M.E.  1989–1990. 
Manual of North American Grasses. Newsletters 1–3.

78.  Barkworth, M.E. and K.M. Capels.  1990–2002. 
Manual of North American Grasses. Newsletters 4–12.

Grant support

Unless noted otherwise, I was the principal investigator on
the following grants.  Those marked with
an asterisk are active; those that are underlined were for graduate student
research projects. 

2010-2011       Utah Agricultural Experiment Station.
Creating Access to the three herbaria of Utah State University. Utah
Agricultural Experiment Station. $20,000.

2009 – 2010.   National Parks Service. Flora of the Green
River in Dinosaur National Monument. $20,000.

2009 – 2010    National Park Zervice. Curation of plant
specimens that were frequented by bees collected during a survey of the bee
fauna of Zion National Park. $5,000.

2006 – 2007    USU-ADVANCE. To organize a collaboration of Intermountain Region
herbaria. $7,000.

2005 – 2007    U.S.D.A. Agricultural Research Service – Forage and Range
Research Laboratory, for completion of volume 24 of the Flora of North America. $359,234

2005 – 2007    National Science Foundation: Completion of volume 24 of
the Flora of North America series. $150,000.

2005                Flora of North America Association. For work
on volume 24. $20,000.

2004                National Fish and Wildlife Foundation. Manual
of Grasses for North America
: editorial work. $29,000.

2004                National Park Service: for illustrating volume
24 of the Flora of North America. $10,000.

2004                Flora of North America Association: $50,000,
for work on volume 24.

2003 – 2004    U.S.D.A. - Natural Resources Conservation Service: for
volume 24, making it available on the Web. $7000.

2003 – 2004    Bureau of Land Management, for Plant Information Network 2.
$9000.

2003 – 2004    U.S. – Armenia Research Foundation: Grasses of Armenia,
(E. Gabrielan co-PI). This funded travel by Frank Smith and myself to Armenia.
$5700.

2003 – 2004    Bureau of Land Management: Flora of North America, volume 24. $10,000.

2003                Flora of North America Association, for work on Flora
of North America volume 24. $50,000

2002                U.S.D.A. - Natural Resources Conservation Service:
 Manual of Grasses for North America,
$15,000.

2002                Bureau
of Land Management:
Manual of Grasses for North America, $20,000         

2001 – 2003    National Park Service: Manual of Grasses for North
America,
$40,000        

2001                Flora North America Organization. Manual of
Grasses for North America.
$20,000.

2000 – 2003    Utah Agricultural Experiment Station. Manual of Grasses
for North America.
$18,000 (adjusted each year – ballpark is $6000 per
year).

2000 – 2002    U.S.D.A. - Animal Plant Health Inspection Service; Manual
of Grasses for North America.
$30,000

2000                Flora North America Organization; Manual of
Grasses for North America.
$43,000.

2000                Bureau of Land Management: Manual of Grasses
for North America
. $10,000

2000 – 2001    U.S.D.A. - Natural Resources Conservation Service: Manual
of Grasses for North America.
$28,000

2000 – 2007    U.S. Forest Service. Manual of Grasses for North America. $175,000

1997                U.S. Forest Service (Umatilla National Forest):
Potential vegetation of the North Fork John Day Wilderness area.

1996                U.S. Forest Service (Payette National Forest):
Reproductive biology of Saxifraga
bryophora
var. tobiasieae.

1995 – 1997    Utah State University Faculty Assistance Center: Resources
for learning basic skills in plant taxonomy $10,000

1995 – 1996    National Science Foundation: Upgrade of herbarium
facilities. $99,970

1992 – 2002    Utah Agricultural Experiment Station: Illustrations for
Manual or North American Grasses. 
$18,750.

1992 – 1997    U.S. Department of AgricultureAgriculture
Research Service
: Manual of Grasses for North America.  $113,000

1990 – 1991    Soil Conservation Service: Distributional data from the
Intermountain Herbarium.  $3,000

1988                United States Department of Agriculture-Forage
and Range Research Laboratory: Morphological comparison of Pascopyrum smithii and Elymus
lanceolatus.
$1,000.

1987 – 1990    International Board of Plant Genetic Resources: Systematic
studies in Thinopyrum.  $27,000.

1987 – 1990    International Board of Plant Genetic Resources: DNA
hybridization studies in diploid Triticeae. 
$10,000.

1987 – 1992    United States Department of Agriculture: Manual of North American
Grasses.  $60,000.

1986                Smithsonian Institution: Systematic studies of Indian
and Pakistani Stipeae.  $5,684.

1985 – 1987    Smithsonian Institution: 
International symposium on grass systematics and evolution.  $85,000 (co-PI).

1982 – 1985    Utah Agricultural Experiment Station: A photographic slide
collection and computer-based catalog.  
$7,000.00

1981 – 1984    National Science Foundation: Systematics of Stipa in North America.  $
60,015.00.

1980 – 1989    Utah Agricultural Experiment Station: Taxonomic studies in the Triticeae.  $54,201

Web sites

----

Plants of South Asia: currently http://usuherbarium.usu.edu/pakistan/, to become http://southasianplants.net.
This site is designed to present specimen- and image-based records of plant
occurrences. It uses Symbiota (http://symbiota.org) software.

Grasses: http://grassesareglorious.net.
This site will present information specimen- and imaged-based records of
occurrences for grasses. The initial focus will be on North American grasses
and will use information obtained when developing Barkworth et al. (2003,
2007a) and their derivative volumes. It will be expanded, as resources permit,
to reflect new information, differing taxonomic positions, and maps plus
information from other resources and other part of the world. It uses Symbiota
(http://symbiota.org)
software.

Intermountain Herbarium Website. http://www.herbarium.usu.edu/.
This site presents information about the herbarium and has subdirectories for
my major research projects, the classes that I teach, information for the
general public, and some general resources. It needs a major “spring cleaning”.

----

Teaching

Courses

I shall be retiring from teaching at Utah State University
on June 30, 2012. The four courses listed are those I have been teaching for
the last two years.

Biology
4420                (3 cr)    Wtr      Plant
taxonomy. 

Biology
2410                (2 cr)    Smr      Plants
and fungi in the field.  

Biology
1020                (1 cr)    Fall       Discovering
Biology

Biology
1760                (1 cr)    Fall       Special
topics: herbarium practice

In the past, I have taught courses in agrostology, advanced
plant systematics, and numerical taxonomy.

----

In retirement, I plan to create online offerings that will
be available in two or three different modes: open access and free, no
requirements; open access and low cost with certificate of accomplishment for
satisfactory performance which will be determined by assignments and an oral
examination, possibly via Skype; as a “for credit” class or part of a for
credit class of Utah State University, at standard university cost.

----

Graduate students

1999.   A.M. Kelly,
M.S.  Thesis: Predictively mapping the
plant associations of the North Fork John Day Wilderness in northeastern Oregon
using classification tree modeling. Published as Kelly, A., D.C. Powell,
and R.A. Riggs. 2005. Predicting Potential Natural Vegetation in an Interior
Northwest Landscape Using Classification Tree Modeling and a GIS. Western
Journal of Applied Forestry 20: 117–127.

1999. K.
Pierson. M.S.            Thesis: The
reproductive biology ad edaphic characteristics of a rare, gynodioecious
saxifrage: Saxifraga bryophora var. tobiasiae (Saxifragaceae).

1991. P.S.
Hoge., M.S.            Thesis: Biosystematics of seven species of Stipa from the southwestern United
States and northern Mexico.

1990, J.K. Jarvie, Ph.D.           Dissertation: The
biosystematics of Thinopyrum (Triticeae: Poaceae).

1984. T. Stockton,
M.S.          Thesis: Xylorhiza glabriuscula; stabilized hybrid derivative or
evolutionary intermediate?

1983. E.E. Neely,
M.S.            Thesis: A comparison of
the vegetation on soils derived from dolomite and quartzite in the Bear River
Range, Utah and Idaho.

1983. R.J. Atkins,
M.S.            Thesis: A taxonomic
study of Leymus ambiguus and L. salinae (Poaceae).

Workshops

10/01/10          Digitizing herbaria at College of Eastern Utah.

06/15/07          Presentation on Grasses, the world’s most successful
plants
to participants in a
workshop for faculty in plant conservation sponsored by the University of
Peshawar Botanic Garden.

11/09/06          Herbarium sponsored workshop on Using interactive
identification keys
. With Tim Jones and Pedro Lake.

03/27/04          Herbarium sponsored workshops on Sedges, offered by
Sherel Goodrich and Sagebrush, offered by Al Winward, both employees of
the USFS. Michael Piep. co-leader..

03/05/04          Herbarium
sponsored workshop on Grasses 2, a workshop for professionals.

01/31/04          Herbarium sponsored workshop on Grasses
1
, a workshop for non-botanists. Attended by 19 individuals. Michael Piep,
co-leader.

Service

Herbarium

My primary goals as Director of the Intermountain Herbarium
are a) to develop the herbarium’s value as a regional resource for information
about plants and fungi in the intermountain region; b) develop its financial
resources; and c) expand the collection.

Regional resource:
I have three activities in this area: Digitizing the collection, offering
workshops, and creating online identification resources. 

Digitization: I initiated databasing specimen data in
1988; georeferencing, primarily of grasses, began in the 1990s; and imaging
after the purchase of equipment in 2009. In 2010, the herbarium began making
its specimen information available through four taxonomically focused websites:
intermountainbiot.org (vascular plants)., mycoportal.org (fungi), nalichens.org
(lichens), and bryoportal.net (bryophytes). It is not, at present, feasible to
have a single portal providing access to all our specimen records.

The herbarium’s workflows have been, and will continue to
be, modified to take advantage of ongoing technological developments that
increase increased efficiency in specimen digitization. Among these
developments is a module that collectors can use for recording collection
information and generating labels. Data recorded in this module can be pulled
directly into herbarium databases when the specimens are deposited, eliminating
the retyping of information that has to take place with labels created by other
means. Consequently, persuading collectors to use the tool has become a
priority.

Workshops. The
herbarium has sponsored a variety of workshops. These were initially well
attended but attendance at recent workshops has been very limited. Because the
benefits from offering them are limited, it seems better to focus on developing
online resources and then using these to help attract greater interest, perhaps
in conjunction with a certificate program.

Online identification
resources.
There have been major changes in the taxonomy of plants and
fungi during the past 20 years, changes that are not currently reflected in
affordable print publications. Providing illustrated online identification
resources will benefit a wide array of user groups.

Discretionary Funding. Establishing ongoing discretionary funding for the herbarium is a
major challenge. Royalties from the grass publications I have edited generate a
significant sum. My hope is that the online identification resources can also
be used to develop hard-copy publications that people will generate royalties.
In the past, the herbarium has sold notecards and prints but the sales do not
justify the investment required.

Expand the collection. Collection growth has been
rather slow in the last two decades, usually fewer than 3000 specimens per year
and mostly through donations and exchange. I plan to start an active regional
collecting program, both to strengthen the collection and my knowledge of the
region’s flora. This last will also improve the herbarium’s reputation as a
center of regional activity.

2012

  • Participated in a workshop sponsored by iDigBio
    on the digitization of herbaria.
  • Led annual meeting of the US Virtual Herbarium
    project.
  • Submitted a request for funding to support
    imaging and digitization of Utah State University’s two small herbaria, USUUB
    and PRI.
  • Revised the imaging workflow, resulting in
    speedier movement of specimen images to the regional network and placing the
    herbarium in a position to take advantage of remote data entry and OCR assisted
    data entry.
  • Encouraged offering of a workshop on
    georeferencing by the Natural History Museum of Utah. The museum offers easy
    access via air and is more central than Utah State University.

2011

  • Continued development of effective workflow for
    digitizing vascular plant specimens with the help of funding from the Utah
    Agricultural Experiment Station. In mid-July, about 9000 images were added to
    the regional web site. Of these, approximately 3000 were connected to records
    already made available to the web site. This fall we shall focus on entering
    the data for the remaining specimens.
  • Helped Snow College
    (EPHR) and the College of Eastern Utah (PRI) become contributors to the
    regional web site.

2010

  • In connection with
    grants for digitizing the herbarium, initiated the reorganization of the Asteraceae so that it reflects current
    treatments. As part of this activity, specimens were annotated, repaired, and
    barcoded.
  • Worked with Ed Gilbert
    in establishing a regional web site, http://intermountainbiota.org.

2009

  • Placed interactive, directed choice keys for
    plants in the Intermountain Region on the herbarium’s web site. These keys
    reflect current taxonomic treatments.
  • Visited Ephraim State College to talk with the
    botanist there about integrating its herbarium into the proposed virtual
    regional herbarium.
  • Began barcoding and updating nomenclature on
    specimens of Asteraceae. This is of
    general benefit but the choice of family reflects the decision by regional
    herbaria to emphasize Asteraceae. The
    activity is included in the NSF proposal that was submitted in July 2009, but
    anything we can do in this regard is beneficial.

Past

  • Initiated the development of most records on
    herbarium activities that are included in the annual report.
  • Developed the initial relational database used
    by the Intermountain Herbarium.  This
    included several authority files, facilities for printing regular herbarium
    labels and annotation labels, and provision for cataloging the slide
    collection.  The database used Alpha
    Software.  The herbarium has since switched
    to a different system – and will be switching again in 2010.
  • Increased the number and variety of individuals
    using the herbarium. It is still not used as much as it should be, but usage
    has increased.
  • Developed Web pages for the herbarium that
    provide access to a variety of different kinds of information, including some
    that is of interest to taxonomists working with the two groups of grasses of
    particular interest to me, the Stipeae
    and Triticeae
  • Made information being developed for the Flora of North America grass volumes
    available over the Web at http://herbarium.usu.edu/webmanual.  We have already updated, amended, and added
    notes to some of the information in the published volume.
  • Initiated the selling prints of herbarium
    specimens and note cards that make use of illustrations prepared for the grass
    project to help provide operating funds for the herbarium.
  • Sponsored the development of free choice interactive
    keys and accompanying fact sheets for grasses and sedges. The Carex resources, and others developed by
    T.M. Jones, will soon be deleted because he is now a graduate student at
    Louisiana State University and regularly updates the resources that he makes
    available.
  • Initiated posting of data to the Global
    Biodiversity Information facility, becoming the first herbarium in the
    intermountain region to do so.
  • Initiated creation of interactive keys and
    descriptive pages focusing on the Triticeae and the plants of the Intermountain
    Region (this has not gone far as yet).

Roles in professional organizations

2012                Convenor,
Special Committee on Registration of plant and algal names (including fossils)
for the General Committee of the International Association of Plant
Taxonomists.

                        Conducted
the second annual survey of digitization in US herbaria.

2011 - 2013     Chair,
International organizing committee for the Seventh International Triticeae Symposium which will be held
in Chengdu, China in 2013.

2011                Initiated
a survey of herbaria in the US in order to obtain current information on their
status. As of August 25, responses had been obtained from over 40% of the 604
herbaria known to exist in the US. Importantly, it has identified herbaria that
have been closed as well as updated information for many herbaria and collected
new information on digitization.

2011 - present  Member, Advisory Board, Botanical Garden of the University of Swat,
Pakistan.

2010-2011       Chair of the Task Force to develop an
organization structure for the US Virtual Herbarium Project. A revision of the
task force’s proposal was approved in January, 2011.

2009                Chaired the annual meeting of
the US National Virtual Herbarium project.

                        Member of International
Review Panel for the herbarium of the University of New England, Armidale,
Australia.

2009 –Now        Co-chair, US Virtual Herbarium
Initiative. The goal of this initiative is to accelerate digitization of US herbaria
in a manner that encourages participation by all herbaria, including small
herbaria, in developing a national digital herbarium. This will help ensure
that all students entering systematic botany learn to record, contribute, and
use specimen based digital information

2008                Created the Coordinating
Committee for development of a US National Virtual Herbarium. Chaired the first
annual meeting.

2007 -2010      Member at large, American Society of Plant
Taxonomists.

2007                Co-organizer of a colloquium on Integrating systematics at the annual
meeting of the American Society of Plant Taxonomists.

2007 – 20011  Member, Committee on Electronic Publication,
International Association of Plant Taxonomists.

2005
– 2008    Member, Building Advisory
Committee, University of Peshawar Botanic Garden

2004                Vision Committee, Herbarium
Network Organization; Member,

2004                Utah Native Plant Society; Board
member.

2003                Environmental
Task Force, University and the Environment; Member.

2001 – 2003    Education and outreach program, American
Society of Plant Taxonomists; Member.

1997 ­– 2000    Member, Editorial Board, Northwest
Scientific Association: Member, Editorial Board.

1999 – 2006    Flora of North America, Editorial Management
Committee; Member.

1995 – 2008    International Triticeae Consortium: Secretary/Treasurer

1988 – 1992
   American Association of University
Professors (USU Chapter): Treasurer

Attendance and presentations at professional meetings

2012    Pakistan
Botanical Garden Network.
SYMBIOTA: Open Source Software for developing specimen-based
floristic resources for Research and Education. (Presented remotely).

            Annual
meeting of the Botanical Society of
America, Columbus, Ohio. Grasses of
North America, a new web site.

2011    International
Botanical Congress
, Melbourne, Australia. Online interactive pathway
(directed) keys: an effective tool for serving today's students while building
resources for tomorrow's students.

Annual meeting of the Botanical Society of
America,
St. Louis, Missouri. North American Meliceae (Poaceae): morphology, geography, and phylogeny.

SYMBIOTA: Providing
tools for public engagement with collection information
. With Ed Gilbert.

Co-chaired (with Zack
Murrill). US Virtual Herbarium
Project; Progress made, goals to achieve
.

2010    Annual
meeting of the Botanical Society of America,
Providence, Rhode Island.

            Digitizing lichens, fungi, and
bryophytes: exploring some options. Poster presentation. Authors: N. D. Stock,
M.B. Piep, and M.E. Barkworth.

2009    Annual
meeting of the Australian Society of Systematic Botany
, Armidale,
Australia.

            The Triticeae in Australasia. Oral presentation.

2009    Annual
meeting of the Botanical Society of America.
Snowbird, Utah.

            Annual meeting of the US Virtual
Herbarium project. Chaired.

            Wandering in a field of daisies.
Oral presentation.

2009    Sixth
International Triticeae Symposium

            Identification of genomic genera.

2009    Meeting
of the North American Nodes to the Global Biodiversity Information Facility
.
Washington, D.C. April 22-24.  Invited
participant.

2008    Fourth
International Symposium on Grass Systematics and Evolution. (Copenhagen,
Denmark).

            Diversification
in the Stipeae and Triticeae. Keynote addres. With  M.O. Arriaga, W-L. Chen, J. Chiapella, C.
Garrard, S.W.L. Jacobs, M. Olanova, P. Shavrova, P. Peñailillo, F.
Valdés-Reyna, and F. Vazquez.

The Triticeae in Australasia. Barkworth, M.E.,
J. Rollo, and S.W.L. Jacobs. Oral presentation.

2008    Botanical
Society of America (Vancouver, Canada).

Building a national
virtual herbarium through collaboration. Workshop.

The Stipeae in Australia and North America:
Observations from the field. Oral presentation.

T          he WDC012 Committee and developing a
single portion to information in US herbaria. Oral presentation, with A.
Liston, M. Mayfield, J. Pollock, and Z.E. Murrell.

2007    Botanical
Society of America.

            Generic delimitation in South
American Stipeae. Oral presentation.
With Mirta Arriaga, J. Smith, J. Valdes-Reyna, S.W.L. Jacobs, and S. Bushman.         

Developing a
national atlas. Oral presentation during colloquium.

2006.   Botanical
Society of America
. No presentation.

            St.
Louis Botanical Garden Annual Symposium
.

            Sponsored
demonstration of an interactive key to Carex
(see http://utc.usu.edu/keys/).

            Consortia
of Intermountain Herbaria
.

Organized, with
Michael Piep, meeting for participants at Utah State University. Highly
productive discussion of ways in which we could collaborate. Led to development
of two grant proposals to the National Science Foundation, one from Wilson of
UVSC, one from Barkworth.

2005    Smithsonian
Botanical Symposium
, “The future of floras: new frameworks, new
technologies, new uses”. No presentation..

2005    Fifth
International Triticeae Symposium
.
See non-refereed papers for presentations.

2005    International
Botanical Congress
. “Bridging the Internet Divide”. Poster presentation
with S.-J. Lim. 

2004    Society
for Range Management
. No presentation – promotion of grass volume and
solicitation of additional funding.

2004    American
Society for Plant taxonomists
. Presented paper and sponsored table where
the herbarium sold prints and note cards and made its activities known.

2004    Herbarium
Network Meeting
: Invited to participate in workshop on development of a
herbarium network. One of 20 invitees from the U.S.A.

2003    Society
for Range Management
. No presentation – promotion of grass volume and
solicitation of additional funding.

2003    Third
International Monocot Symposium
. From regional to global without being
colonial: expanding the Web site for the Manual of North American Grasses.
Poster presentation to the. With Jesús Valdes Reyna.

2003    XXIX
Jornadas Argentinas de Botánica.
Development of a Global Grass Resources
Network. With Seung Jin Lim, Bryan Heidorn, Mirta Arriaga, and Jesus Valdes
Reyna. Invited talk.

2002    American
Society of Plant Taxonomists.
Disentangling distributional information: Echinochloa
crusgalli
and E. muricata.

2001    Fourth
international Triticeae Symposium
.
Voucher Specimens: an essential element in scientific research (with S.W.L.
Jacobs). Oral presentation.

            The Triticeae in the Americas (with M.O. Arriaga).  Poster presentation. 

2000    Northwest
Scientific Association
. The Manual of Grasses for North America: Current
status and prognosis. (with K.M. Capels and L.A. Vorobik). Poster presentation.

2000    Society
of Range Management.
The Manual of Grasses for North America: Current
status and prognosis. (with K.M. Capels and L.A. Vorobik). Poster presentation.

2000    American
Society of Plant taxonomists
. The Manual of Grasses for North America:
Progress Report. (with K.M. Capels and L.A. Vorobik). Poster presentation.

1999    International
Botanical Congress.
A Grass Manual for North America: Concept, progress,
and prognosis. (with K.M. Capels and L.A. Vorobik). Poster presentation.

1999    Symposium
on Eastern Native Grasses, sponsored by the USDA-ARS.
A Grass Manual for
North America: Concept, progress, and prognosis. (with K.M. Capels and L.A.
Vorobik). Poster presentation.

1998    Third
International Symposium on Grass Systematics and Evolution.
 Grasses in North America: A biogeographic
perspective.
(with K.M. Capels) [Invited talk]    

Third International
Symposium on Grass Systematics and Evolution. 
Changing perceptions of the Triticeae.
[Invited talk].

1998    Third
International Triticeae Symposium
:
The Triticeae in North America

1997    Northwest
Scientific Association
: The Virtual Herbarium: progress report and
workshop

1996    American
Society of Plant taxonomists
: A Virtual Herbarium: what purpose should
it serve and how should it be developed

1994    Northwest
Science Association
: Distinguishing Elymus
lanceolatus
from Pascopyrum smithii

1994    Second
International Triticeae Symposium
:
The Elymus trachycaulus complex:
more questions than answers

1991       
California
Botanical Society
: Old flames, new names (Invited speaker)

1991    Northwest
Scientific Association
: Five years of herbarium databasing: lessons
learned the hard way

1990    Plant
Molecular Biology Seminar Series, Montana State University
: Evolution in
the Triticeae: A classical
systematist's view

1990    American
Society of Plant Taxonomists
: Genera of North American Triticeae

1988    Botanical
Social of America, western section
: Cladistics: Current Boondogle?

1988    American
Society of Plant Taxonomists
: Anatomical variation in Thinopyrum and its allies (Poster,
with J. Jarvie)

1988    American
Society of Plant Taxonomists
: GEDS: A geographic and ecological database
for plants
(Poster, with J. Choules)

1988    Annual
Symposium of the Agricultural Research Service, Beltsville
: Manual of
North American Grasses: Progress Report
(Poster)

1987    American
Society of Plant Taxonomists
: Plant distribution databases: content and
scope

1986    International
Symposium on Grass Systematics & Evolution
: Identification and
relationships of monophyletic groups in the Stipeae

1986    American
Institute of Biological Sciences
. [No presentation].

1984    Tinker
Conference on Biogeography of Mesoamerica
: Piptochaetium in North and Mesoamerica: phytogeographical
Implications

1984    American
Society of Plant Taxonomists
: The Generic Affiliation of Stipa sect. Stephanostipa

1983
   American
Association for the Advancement of Science, western section
: Genomically
based genera in North American Triticeae

1983
   American
Society of Plant Taxonomists
: Leymus
Hochst. in North America

1983
   Botanical
Society of America, western section
: Adaptive trends in the evolution of
North American Stipeae

1983    American
Society of Plant Taxonomists
: Adaptive Trends in the Evolution of North
American Stipeae

1982    Department
of Biology, University of Ottawa
: Parallel evolution and taxonomic
confusion in the Stipeae

1982
   Botanical
Society of America, western section
: The generic affiliation of Oryzopsis kingii

1981
   International
Botanical Congress
: Ptilagrostis
in North America

1981    International
Botanical Congress
: Leaf cross‑sections and the taxonomy of the Stipeae in North America

1980    American
Society of Plant Taxonomists
: The Stipeae
in North America

1979
   American
Society of Plant Taxonomists
: Intraspecific taxa in Stipa nelsonii Scribner (= Stipa
columbiana sensu
A. Hitchcock)

1979    Canadian
Botanical Association
: Evolutionary and taxonomic relationships in North
American Stipeae

1978
   Canadian
Botanical Association
: Paleal morphology and the taxonomy of  Poa

1977
   American
Society of Plant Taxonomists
A
taxonomic study of large‑glumed species of 
Stipa in Canada

1974
   Northwest
Science Association
Intraspecific
variation in Brodiaea douglasii
Watson

Other professional activities

2007          Organized Colloquium for annual
meeting of American Society of Plant

                  Taxonomists

                  Participated in a workshop for faculty in plant
conservation sponsored by the University of Peshawar Botanical garden and
participated in discussions concerning a graduate curriculum in plant
systematics.

2005          Discussions and field work in
Pakistan; Member,

                  Building Advisory Committee,
University of Peshawar Botanic Garden.

2004          Fieldwork in Sweden and Denmark

2004          Fieldwork and herbarium study Armenia

2003          Field and herbarium study work in
Argentina

2003          Fieldwork in Armenia

2005          Fieldwork in California

2000          Botanical Society of America: Member, judging
committee for student                                 presentations.

1999          Fieldwork on North American Stipeae

1998          Agricultural University of Sweden,
Svalöv: Designated opponent,                                         dissertation
defense of Claus Baden.

1998          Field work in Australia with Dr.
S.W.L. Jacobs, Royal Botanic Garden

                  Sydney. 

1988          International Board of Plant Genetic
Resources Training Program:                                        member
of review panel.

University committees

?-9/10              Ombudsman for
the College of Science

09/99-09/07                 Scholarly Publications Committee,
Utah State University; Member.

9/96 – 12/02    Tenure Committee, C. von Dohlen, Biology Department, U.S.U.; Member.

9/96 – 6/98      Graduate Program
Committee, Biology Department; appointed member

9/96 – 3/97      Vertebrate Systematist,
Biology Department; appointed member

7/95 – 6/98      Adjunct Status Committee,
Biology Department, U.S.U; appointed member

7/94 – 6/97      Athletic Council, Utah
State University; member elected by Senate

9/90 – 6/91      Tenure Committee, F.
Messina, Biology Department, U.S.U.; appointed member

7/90 – 6/93      Athletic Council, Utah
State University; elected by senate

10/89 – 6/90                Term limits
committee, University Senate; appointed to chair by senate

9/86 – 6/90      Arboretum Committee, Utah
State University; appointed member

7/86 – 6/89      Honorary Degree Committee,
Utah State University; appointed by senate

9/86 – 6/88      Graduate Program
Committee, Biology Department; appointed chair

9/85 – 8/86      Graduate Program
Committee, Biology Department; appointed member

Other

Reviewed articles, when requested, from American Journal of
Botany, Annals of Missouri Botany Garden, Australian Journal of Botany, Economic
Botany, Genome, Great Basin Naturalist, Madroño, Northwest Science, Novon,
Systematic Botany, Telopea.  I have also
reviewed proposals for the National Science Foundation, the Idaho Research
Board, the Montana Research Board, and the National Geographic Society.

Memberships

American Association for the Advancement of Sciences;
American Institute of Biological Sciences; American Society of Plant
Taxonomists; Association of Natural History Collections; Botanical Society of
America; California Botanical Society; International Association of Plant
Taxonomists; International Triticeae
Consortium; Northwest Science Association; Taxonomic Database Working group,
Utah Native Plant Society.

Occasionally relevant skills

  • Moderate reading ability in Latin, French,
    Spanish, and minimal ability in Russian. 
    Speaking and writing ability in these languages is essentially nil.
  • Basic ability with HTML language and the
    following software: Photoshop, Illustrator, InDesign, Dreamweaver, ArcView,
    Alpha 5; Microsoft Office components (minimal knowledge of Access), Microsoft
    Publisher.

Personal

BIRTHPLACE: Marlborough, Wiltshire, England

CITIZENSHIP: Canada           U.S.
VISA STATUS: Permanent

STATEMENTS OF INTEREST

Research

My research interest is plant systematics, with a particular
interest in grasses. At present, I am engaged in work on the Triticeae. This is an important tribe of
grasses because it includes three major cereals (wheat, barley, rye) plus ~350
additional species many of which are important forage species. Its taxonomic
treatment is complicated by the low barriers to hybridization and rampant
polyploidy within the tribe. This is exacerbated by the widespread distribution
of the tribe (it is represented in almost all temperate regions of the world)
and the discrepancy between the detailed information about the cereal species
(e.g., the chromosomal location of the genes controlling awn development) and
the minimal information available for many othe species, some of which are known
from only a few specimens. The taxonomic problems in the tribe involve both
circumscription of its perennial genera and that of most of its species. My
primary interest is in the generic problems but to develop good generic
descriptions, one needs good species descriptions. Moreover, because the tribe
is almost global in extent and includes many species that have been introduced
beyond their native range, these descriptions need to be web accessible. I have
posted various resources for those working on the Triticeae to the web at http://herbarium.usu.edu/triticeae/.
At present, I am working on a revision of the Australasian species. This work
developed from observations made during my sabbatical. [Two publications out]

In 2010, I initiated a study of the phylogeny of Melica in collaboration with Dr. B.S.
Bushman (USDA_ARS). This is a relatively small tribe at the base of the Pooideae that is found in temperate
regions of Eurasia, North America, and South America. It is almost completely
absent from Australia. Previously, Bushman and I had worked on the other large
genus in the tribe, Glyceria. At present, we are developing a
chloroplast phylogeny for North American members of Melica and another, primarily North American member of the tribe, Pleuropogon. In addition, I am creating
species pages for the individual species of Melica
 by translating published accounts.

In 2008, I developed a major data set concerning the
distribution of morphological characters in the Stipeae that will be written up in 2011. I started writing it up in
2009 but had to give other tasks priority. The paper will offer a new way to
look at and synthesize morphological data on a global level. My earlier goal
had been to work on the phylogeny of the tribe and I collected specimens of the
Australian species for this purpose during my sabbatical. That summer, I was
made aware that another, well funded, group was working on the tribe; there was
no interest in collaborating.

For 20 years, my responsibility for the collaborative
development of a comprehensive treatment of the grasses of North America, Flora of North America (FNA) volumes 24
and 25, overshadowed my research activities. The two volumes were published in
2007 and 2003, respectively. The information in them was then cut and condensed
to create a single volume treatment, the Manual
of Grasses for North America
which was also published in 2007. This yielded generated about $10,000
in its first year, some of which was used to pay for a second printing, some
for creation of Grasses of the
Intermountain Region
(see next paragraph). The remainder has been used to
fund my research and support the herbarium. I also, as time permits, update the
pages on the Web that were created initially from the FNA volumes,

Teaching

I teach three “normal” classes and offer one “Special
topics” class. The three normal classes are Plant taxonomy 1, Plants and fungi
in the field, and Discovering biology. Plant taxonomy 1 emphasizes plant
identification but does not require learning of family characteristics, as is
the vase of Plant taxonomy 2. The two aspects used to be in a single course but
were split at the suggestion of some students who pointed out that offering
plant taxonomy 2 in fall would enable students to use their skill in identification
over the summer, making it easier to appreciate the families. In northern Utah,
there are very few native or established plants available for study until the
last two weeks of “spring” semester, the time when Plant taxonomy is taught.
This time slot was chosen because the students who are required to take plant
taxonomy also have to take many other field courses in the fall.

Plants and fungi in the field, BIO 2400, is a 2-credit
course that attracts about 20 students. Students appreciate the high proportion
of time spent in the field and the fact that it is offered during the gap
between the regular academic session and the standard summer session. The
course includes three short writing assignments. Dr. B.R. Kropp covers the
fungal aspects of the course.

Discovering Biology is a lab science course that is taken
primarily by students in exercise science and nursing. The students spend most
sessions conducting small experiments for which they then have to write
reports. In my version of the class, all the experiments involve plants or
fungi.

 The “Special Topics”
course is herbarium studies. This enables students to gain experience working
in a major research herbarium, as part of which they learn about plant
taxonomy. It is a hands-on course, but what the students do depends on their
background and interest.

Herbarium

As Director of the herbarium, I am trying to improve its
financial health, visibility, and use. This has led me in two directions. One
is publishing books, with monies earned going to my research and the
herbarium’s budget. The other is digitization of herbaria and development of
portals to the holdings of multiple herbaria. Increasing use falls primarily to
the Assistant Curator who assists users and offers workshops.

The two FNA grass volumes have generated about $800 per year
since they were first published. The next volume published, the Manual of grasses for North America,
yielded over $10,000 in its first year and, because it is being adopted as a
text book in agrostology classes, continues to provide about $2400 per year. It
is in its second printing. The third book, Grasses
of the Intermountain Region
, was published in fall 2009. It yielded $4000
in its first year. I have no plans for publishing another book along these
lines but may bring out a second edition of the Manual in 2012. This will reflect taxonomic and distributional
changes made since publication of the first volume.

For around twenty years I have been working to make the
herbarium database available on line. We provide over 90,000 records to the
Global Biodiversity Information Facility (GBIF), but we have databased over
100,000. Unfortunately, the server that held the database and served the data
crashed in August 2010 and the new server, which was with a combination of
donations and herbarium funds in April 2010, is still (10 October 2010) not
available for use although it is getting closer. Once it is fully operational,
we shall convert to a new and more powerful database developed by Dr. Leigh
Johnson of Brigham Young University.

In addition to purchasing a server, the herbarium purchased
the equipment needed to start imaging its specimens, again relying primarily on
donations. In June, 2010, we were awarded $20,000 from the Utah Agricultural
Experiment Station for digitizing the collection, the primary goal being to
improve the competitiveness of the NSF application that was submitted on July
23, 2010. Because the new server was not available before submission, we were
unable to make as much progress as we had hoped. We did, however, hold the
proposed workshop at the College of Eastern Utah and have loaned the equipment
to them so that they can image their specimens.

http://herbarium.usu.edu/mary.html

Taxonomic name: 
Festuca (World Grasses), Glyceria (World Grasses), Hakonechloa (World Grasses), Hesperostipa (World Grasses), Heteropogon (World Grasses), Hilaria (World Grasses), Hordeum (World Grasses), Hyparrhenia (World Grasses), Leymus (World Grasses), Macrochloa (World Grasses), Melica (World Grasses), Miscanthus (World Grasses), Munroa (World Grasses), Nassella (World Grasses), Oryza (World Grasses), Oryzopsis (World Grasses), Panicum (World Grasses), Pascopyrum (World Grasses), Phalaris (World Grasses), Phleum (World Grasses), Phyllostachys (World Grasses), Piptatherum (World Grasses), Piptochaetium (World Grasses), Pleuropogon (World Grasses), Polypogon (World Grasses), Polytrias (World Grasses), Pseudoroegneria (World Grasses), Ptilagrostis (World Grasses), Reimarochloa (World Grasses), Scolochloa (World Grasses), Secale (World Grasses), Setaria (World Grasses), Sorghum (World Grasses), Spartina (World Grasses), Stipa (World Grasses), Themeda (World Grasses), Thinopyrum (World Grasses), Tripsacum (World Grasses), Trisetum (World Grasses), Enteropogon (World Grasses), Elymus (World Grasses), Ehrharta (World Grasses), Distichlis (World Grasses), Elytrigia (World Grasses), Elionurus (World Grasses), Dichanthium (World Grasses), Deschampsia (World Grasses), Cynodon (World Grasses), Cymbopogon (World Grasses), Ctenium (World Grasses), Connorochloa (World Grasses), Cladoraphis (World Grasses), Chloris (World Grasses), Bromus (World Grasses), Anthoxanthum (World Grasses), Anthosachne (World Grasses), Ammophila (World Grasses), Amelichloa (World Grasses), Agropyron (World Grasses), Aeluropus (World Grasses), Achnatherum (World Grasses), Leymus ambiguus (World Grasses), Pascopyrum smithii (World Grasses), Pseudoroegneria spicata (World Grasses), Setaria parviflora (World Grasses), Stipa clandestina (World Grasses), Stipa lemmonii (World Grasses), Stipa robusta (World Grasses), Elymus trachycaulus (World Grasses), Elymus calderi (World Grasses), Bromus carinatus (World Grasses), Dinebra (World Grasses)
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith